In the present day, scientists in the Arctic uncover a circular object with a red, white and blue motif. In March 1942, Nazi officer Johann Schmidt (Hugo Weaving) and his men invade Tønsberg, Norway, to steal a mysterious tesseract possessing untold powers. In New York City, Steve Rogers (Chris Evans) is rejected for World War II military duty due to various health and physical issues. While attending an exhibition of future technologies with his friend Bucky Barnes (Sebastian Stan), Rogers again attempts to enlist. Having overheard Rogers' conversation with Barnes about wanting to help in the war, Dr. Abraham Erskine (Stanley Tucci) allows Rogers to enlist. Rogers is recruited as part of a "super-soldier" experiment under Erskine, Colonel Chester Phillips (Tommy Lee Jones) and Peggy Carter (Hayley Atwell). Phillips is unconvinced of Erskine's claims that Rogers is the right person for the procedure but relents after seeing Rogers commit an act of self-sacrificing bravery. The night before the treatment, Erskine reveals to Rogers that Schmidt underwent an imperfect version of the treatment, and suffered side-effects.
In Europe, Schmidt and Dr. Arnim Zola (Toby Jones) successfully harness the energies of the tesseract, intending to use the power to fuel Zola's inventions. Schmidt, having discovered Erskine's location, dispatches an assassin to kill him. In America, Erskine subjects Rogers to the super-soldier treatment, injecting him with a special serum and dosing him with "vita-rays". After Rogers emerges from the experiment taller and muscular, one of the attendees kills Erskine, revealing himself as Schmidt's assassin Heinz Kruger (Richard Armitage). Rogers pursues and captures Kruger but the assassin commits suicide via cyanide capsule before he can be interrogated.
With Erskine's death the super-soldier formula is lost. U.S. Senator Brandt (Michael Brandon) has Rogers tour the nation in a colorful costume as "Captain America" to promote war bonds rather than be confined to a lab while scientists attempt to rediscover Erskine's formula. In Italy 1943, while touring to active servicemen, Rogers learns that Barnes' unit was lost in battle against Schmidt's forces. Refusing to believe Barnes is dead, Rogers mounts a solo rescue attempt with Carter and Howard Stark (Dominic Cooper) flying him behind enemy lines. Rogers infiltrates the fortress belonging to Schmidt's HYDRA organization, freeing Barnes and the other captured soldiers. Rogers confronts Schmidt who reveals his face to be a mask, removing it to display the red-colored, skull-like face that earned him the sobriquet the Red Skull. Schmidt escapes and Rogers returns to base with the freed soldiers.
Rogers recruits Barnes, Dum Dum Dugan (Neal McDonough), Gabe Jones (Derek Luke), Jim Morita (Kenneth Choi), James Montgomery Falsworth (J. J. Feild), and Jacques Dernier (Bruno Ricci) to attack the other known bases belonging to Hydra. Stark provides Rogers with a new outfit and a new, circular shield made of vibranium, which negates large amounts of damage by absorbing vibrations. Rogers and his team attack and destroy all of the known bases. The team later attacks a train carrying Zola. During the assault Zola is captured but Barnes falls from the train and is lost. Using information gathered from Zola, Rogers leads an attack on Schmidt's final base to stop him from using WMDs against American cities. Rogers clambers aboard Schmidt's jet before it takes off, and during the fight with Schmidt, Rogers damages the tesseract's container. Schmidt physically handles the tesseract, causing him to dissolve in a bright light. The tesseract falls to the floor of the plane, melting through the surface and falling to Earth.
Seeing no way to safely land the plane without risking its weapons detonating, Rogers crashes it in the Arctic. Stark later recovers the tesseract from the floor of the ocean but the aircraft and Rogers remain undiscovered. Rogers awakens in a hospital room. Deducing that something is wrong, he flees outside into what is revealed to be present-day Times Square. There Nick Fury (Samuel L. Jackson) informs him he has been "asleep" for nearly 70 years.
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